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单选题
____
A

except

B

besides

C

despite

D

without


参考答案

参考解析
解析:
本句话前面讲到赤潮导致鱼死亡,接着讲赤潮使贝类中毒,可见这个关联词应该是“除了……之外,还有……”,所以正确答案为B。
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